I have both amps and love them both for different reasons. The Special is PURE Class A operation and runs/sounds like a high quality amp of that type. Class A type amps are not very effecient and usually are not very powerful ( the Special is 30/35 watts)
However, the tone available within that 30/35 watts (and the 15 watt and 5 watt single ended modes) is really awesome (read
Special!)
It provides plenty of clean headroom and lead drive for me at most of the smaller clubs I gig. It also is a lot easier to haul than my 2x12 Lonestar (weighs less and is smaller) so I will always grab it unless I'm playing outside or at a large venue. I also have a 2x12 Lonestal extension cab that I have never taken out of the house. Not sorry I bought it, just haven't needed it.
The Lonestar has tons more clean headroom for several big reasons, one it's Class AB operation which makes it more effecient and allows the amp to produce more power (by design). It's also got 100 watts! So it can be set up so it's a long way to the top.
The Special is loud when it's cranked up, the Lonestar is really LOUD when cranked.
The tubes in the Special (EL-84'S) are a different beast than the 6L6's in the Lonestar,and just putting them (EL84's) in a Lonestar using Yellow Jackets will not yield the same results as they would in the Special. The class of operation is different and the Class A operation is what the Special is all about, the EL84's just work especially well in that situation and that's why they are used. You can't make a Lonestar sound like a Special, they are different animals entirely.
The big 6L6 bottles are best known for producing lots of clean power and by nature are "stiffer" that the EL84's and will not break up as early,and when they do, it's a different, harsher power clip. Better to drive the pre-amp to clip and amplify that with
the 6L6's.
The answer is to own one of each when you can, I love them both. My first Mesa amp was a Lonestar Special, at the time I also owned a Twin Reverb. After playing the Special for a while I went down and demo'd the Lonestar. I immediately ordered a new Lonestar in the same color as the Special (I like the color, O.K.),I also ordered the extension cab at the same time.
I swapped the Twin on the Lonestar. When my new Lonestar came in, I sold my Special to a friend for $1300. In about two weeks I realized what a mistake selling the Special was. I ordered another one and had to wait for 3 months to get it, and then had to pay $1800 for it to boot!
I always learn things the hard way.........
I am not an audio pro so the above is just my opinion/understanding of the difference's in the two amps. But I have tried to give you as much insight as I could. Hope you find it helpful,or at least interesting.
See 'ya, Jim