The difference between the series I RK3 and RK4 boards seems to be just in the reverb circuit.
On the RK4 the reverb return signal from the tank is no longer muted (grounded) and completely killed off when switching, instead the positive feedback on the 2nd opamp is reduced alot, so that the reverb is turned down instead of completely killed. With the RK4 the 1-2 second delay before the reverb is mixed with the dry signal after switching is still there, I don't understand what this circuit revision's for.
This one is a little mundane - also in the reverb circuit (rev. ext. switching bit), a few components were changed/added probably because of problems with external switching systems or pedals damaging the reverb muting transistor. A cap and transient voltage suppressor diode were added to protect the FET muting transistor from voltage spikes caused by switching.