So if it is foot-switchable as well as switchable from the mid pull-pot, it must be actuating an LDR or relay (like with channel switching via its footswitch or pull-pot). If some LDR/Relays are involved, lots of changes could be happening in the amp. That makes it harder to guess about what is going on.
Your added info about the 0.47u cap and the resistor network is very intriguing...but, unfortunately, it doesn't make 100% sense to me. The 0.47u cap is fine...though that's pretty small. Do you know if the cap is on V2A or somewhere else?
Also, what is more confusing to me is your comment about the resistor network. A resistor network makes no sense on the cathode of V2A or of any other cathode, so it must be brought into the circuit somewhere else. The info about the availability of a foot-switch (ie, it being evidence that there are relays/LDRs involved) allows this resistor network to be located practically anywhere. Was there any mention of where it was?
My wild guess is that it is a resistor in parallel to the 680K resistor (to ground) just prior to grid input of V2A. If the 0.47u cap is really on V2A, it's input headroom to mid and upper mids and high frequencies is now really reduced (ie, its output gain is tremendously increased). If you keep the circuit as-is, the full signal strength will really make V2A over-saturate and it'll sound not quite like how most people would prefer. Putting a resistor-to-ground in parallel with the 680K will cut down the signal a little before it reaches V2A. That'll keep the tube from over-saturating (ie, turning to complete mush).
Because of the in-line caps in the main signal path, one could describe the resulting change as reducing some bass while keeping the upper mids intact while keeping V2A from getting too saturated. That kinda sounds like what you described. The trouble with my theory is that it only adds one resistor and not a "resistor network". So, my conjecture could be completely wrong.
All the info you've gotten so far is so juicy. This is so fun. I really appreciate what you've shared. Of course, I'd love even more info or, dare I dream, a schematic of the mods. Man, that'd be sweet.
Thanks for the fun!
Chip