I think I know what the answer will be with the 5751 in V4, it should sound about the same as a 12AX7. Since I have one (may need another to replace it, the getter flash was silver in color on the top of the tube before I tried It in one of the primary stages on a Stiletto Deuce, in a few seconds the flash turned black after I powered up that amp. I believe the tube is still good so will give it a try in V4 this weekend. Some of the long plate 12AX7 will have a similar effect but will not cure the brittle response at higher volumes and you will be limited low gain settings and reduced treble but the presence will definitely add that edge to the grind. I have used Mullard 12AX7 Long plates in V4, V5, and V6. Not bad but no way close to that amazing sound of the JP-2C. JJ ECC803s also similar as it has lower gain factor than the short or medium plate variants. The key to the right tone (in my opinion) is the JAN/Phillips 12AT7 as it adds some warmth to the gain character and reduces the upper harmonics caused by distortion. Mesa 12AX7 sounded bright but much better than the long plate variant of the 12AX7. I have used a 5751 in the V1 position with the Mark V and fell in love with the tube for use with CH1 and CH3 but changed the character of CH2 which is what I generally used for heavy metal style set to crunch voice with the gain dimed, presence just below noon, mids set to about 2pm with the bass line pointing at the mid line on the knobs and treble set to taste. CH2 has that sound but CH3 was sterile. 12AT7 in V4 makes the amp sing with CH3 and does not hinder the performance characteristic that I could tell. Instead, it provided 6 distinct voices in the JP-2C CH2 character (6 voices comes from the three voice toggle switch on the front, the pentode/triode switch doubles that number.
When I first got the Mark V I did not like the 90W character on CH3 so I basically used 45W mode with variac power and I was happy. That in itself led to some other issue as I believe I was pushing the amp harder to get the volume where I wanted it only to find I was killing my power tubes. Ever since then I ran 90W full power all the time and have not had much of an issue with the power tubes red plating but had to reduce treble, gain and presence on CH3 to have something at least usable. V4 change, different story for CH3 and gives you that boogie tone everyone is after, now it is my favorite and that stretches across the board. I still love the JP-2C but I think the change in character on the Mark V with the tube swap has placed this amp on a different pedestal and may rival the JP-2C to some extent. Had I found this magic mojo before I got the JP-2C, I may not have bought it (not true as the JP has features I wanted plus a Class A/B power amp so that was hard to ignore, in other words, I am happy I have both V and JP). Note that change in V4 to the JAN/Phillips 12AT7 does not kill off the cut or brightness of the amp, It is just as bright as the JP-2C. What it does do is regains some of the midrange that is missing in the lower frequencies and rolls off the top end harmonics (takes away the brittleness and sterile tone). I was done with the Mark V before this, was getting ready to sell it and did not expect such a huge improvement. Best of all, it has no effect on CH1 or CH2 unless you change V6 when you use the FXloop set to active. All this talk about tubes makes me want to experiment with my box of 12AX7 variants including the 5751 which does fall into the 12AX7 class due to its gain factor.
When I first got the Mark V I did not like the 90W character on CH3 so I basically used 45W mode with variac power and I was happy. That in itself led to some other issue as I believe I was pushing the amp harder to get the volume where I wanted it only to find I was killing my power tubes. Ever since then I ran 90W full power all the time and have not had much of an issue with the power tubes red plating but had to reduce treble, gain and presence on CH3 to have something at least usable. V4 change, different story for CH3 and gives you that boogie tone everyone is after, now it is my favorite and that stretches across the board. I still love the JP-2C but I think the change in character on the Mark V with the tube swap has placed this amp on a different pedestal and may rival the JP-2C to some extent. Had I found this magic mojo before I got the JP-2C, I may not have bought it (not true as the JP has features I wanted plus a Class A/B power amp so that was hard to ignore, in other words, I am happy I have both V and JP). Note that change in V4 to the JAN/Phillips 12AT7 does not kill off the cut or brightness of the amp, It is just as bright as the JP-2C. What it does do is regains some of the midrange that is missing in the lower frequencies and rolls off the top end harmonics (takes away the brittleness and sterile tone). I was done with the Mark V before this, was getting ready to sell it and did not expect such a huge improvement. Best of all, it has no effect on CH1 or CH2 unless you change V6 when you use the FXloop set to active. All this talk about tubes makes me want to experiment with my box of 12AX7 variants including the 5751 which does fall into the 12AX7 class due to its gain factor.