Just a word of correction here; based on Mesa's description the Lonestar Special is all Class A. Class refers to the bias setting of the amp which (along with some other parameters) determine whether the tube amplifies the positive and negative half of the signal (class A), just one half (Class B), or one half and a bit more (class AB).
Class A amplifiers can be operated single ended since it amplifies both halves of the signal, or they can operate in a push-pull arrangment for higher power.
Class AB amplifiers can only be used in a push pull arrangement so that you get both halves of the waveform.
The Lonestar Special is always class A, the tube biasing is set so that all the power tubes amplify both halves of the signal. in 5W mode it runs one tube single ended, in 15W mode it runs two tubes push-pull, in 30W mode it runs two more tubes (4 total) in parallel push pull.
Tube matching is very important in a class AB amplifier because the designer has selected a preferred crossing point where positive and negative halves of the signal ampifier cross over, and bias level is really a current setting that could very significantly from tube to tube. I'm not sure about the importance in Class A, although I assume it make some difference in absolute maximum power output, but I don't know enough here, so I really can't answer the original question.
good links on Tube Amp Class and Arrangement:
I like Aikens explaination, way more technical.
http://www.aikenamps.com/ClassA.htm
Randall Smith has the best pictures:
http://www.mesaboogie.com/US/Smith/ClassA.htm