V4 is the obvious as it’s for both the send and the return. Intermittent loop jack is another, but since you’ve ruled out those, I think there’s not much else you can do without getting the chassis out and start poking around. You could further narrow it down running the signal from send to another amps return to see does the send provide decent signal. If it does, then problem is between return and V5. Could be wire from pcb to volume coming loose, faulty pot or one of the mute circuit fets.